Orobanche pancicii
Orobanche pancicii G. BECK in Ann. Naturhist. Hofmus. Wien 2: 148, t. 7, fig. 8 - 12. 1887; BECK & SZYSZYLOWICZ, Pl. Cernagoram et Alb. 142. 1888; BECK, Monogr. Orob. 220. 1890; VELENOVSKY, Fl. Bulg. 440. 1891; FORMÁNEK, Zweiter Beitr. Fl. Serb. Maced. Thess. 62. 1895; HAYEK, Fl. Pen. Balc. II: 225. 1929; ROHLENA, in Preslia 20/21: 284. 1941-42; STOJONOV & STEFANOV, Fl. Bulg. 1057. 1948; KITANOV & PENEV, Fl. Vitosha 335. 1963; STOJANOV et al., Fl. Bulg. 2: 996. 1967; BENAC, Fl. Bosn. Herc. 4: 108. 1967; CHATER & WEBB, Fl. Eur. 3: 290. 1972; PARABUTSCHSKI in JOSIFOVIC, Fl. SR Serbije 6: 298. 1974; STRID & TAN, Mtn. Fl. Greece 2: 272. 1991; UHLICH et al., Sommerw.-arten Eur. 107. 1995; DELIPAVLOV in Fl. Reip. Bulg. 10: 310. 1995; PUSCH, Flor. Rundbr. 34 (1): 29 ff. 2000; KIRÁLY et al. in Fl. Pann. 3: 17 ff. 2005; STOYANOV in Proc. IV BBC Sofia 251. 2006; PUSCH in HEGI, Illustr. Fl. Mitteleur. VI, 1A: 47. 2009; DAKSKOBLER et al., in Haquetia 9, 2: 171 ff. 2010
Synonyms:
- Orobanche reticulata f. viscosa G. BECK, Monogr. Orob. 220. 1890 sensu PUSCH, Flor. Rundbr. 34 (1): 37. 2000; FRITSCH in Österr. Bot. Z. 75 (10-12): 223. 1926
Etymology: - The species taxon "pancicii" is dedicated the serbian botanist Josef PANCIC (1814-1888).
Vernacular names:
- Skabiosen-Sommerwurz [german]
- Pančićs Sommerwurz [german]
- Панчичев воловодец [bulgarian]
Description: - Perennial. Stem 25 - 27 cm × (3-) 5 - 8 mm, simple, vigorous, yellowish- to reddish-brownish, sometimes tinged with violet, reddish-brownish glandular. Leaves numerously, dense and imbricate at base of the stem, sparse above, often under the spike absent, anguste lanceolate, 12 - 26 (-30) mm, dense glandulary lanate. Bracts ovate and longe acuminate to ovate-lanceolate, similar to leaves, numerous veined, to 25 mm, dark and dense glandular hairy, equalling or shorter (sometimes longer) than flowers. Inflorescences 6 - 15 × 3 - 3,5 cm, with numerous flowers, ± dense, often lax below, often comose at apex. Calyx 12 - 15 mm, segments brodly ovate at base, unequally bidentate or entire, 5- to 8-veined, teeth shortly triangular, diverge, dark glandulary, 3-veined at base, 1-veined above, middle vein distinctly, equalling the half of corolla to subequalling the corolla tube. Corolla (15-) 20 (-23) mm, erect, pale tawny yellow with purplish veins, densely covered with short, dark purple glandular hairs (especially to the edge); tube broadly cylindrical-campanulate, 6 - 9 mm wide, evenly curved or almost straight in lower 2/3; upper lip slightly emarginate, lobes antrorse; lobes of the lower lip equally or the middle bigger, crenate to almost entire, usually glandular-ciliate. Filaments inserted c. 2 mm from base of corolla tube, shortly hairy at base and in the lower 1/3, above (sub-) glabrous. Anthers brownish in dryness, lanceolate-ovate, whitish pungent. Stigma large, bilobed, yellow to orange-brownish. Style short, glandular hairy. Capsule shortly hairy, c. 9 mm (or more).
Lower taxa:
| 1. | Filaments hairy below, numerous glandular hairy above. (Orobanche pancicii var. typica). | 2. |
| - | Filaments at whole length glabrous or only below very sparse pilose. Style like the ovary numerous glandular hairy. | Orobanche pancicii var. adusta |
| 2. | Bracts as long as the flowers. Calyx segments broadly ovate. Teeth mostly broad lanceolate, shorter than the corolla tube. | 3. |
| - | Bracts longer than the flowers. Calyx teeth elongate, narrow, long acuminate, as long as the corolla tube. | Orobanche pancicii f. longebracteata |
| 3. | Corolla ± dark coloured, covered by numerous dark coloured glandular hairs. | 4. |
| - | Whole plant colured like wax or sulphurous. Glandular hairs at corolla not or hardly coloured. Stigma coloured like wax and the style not tinged with reddish-purple. | Orobanche pancicii f. kerochroa |
| 4. | Flowers large, ca. 20-28mm long. Dorsal line mostly slightly curved. | 5. |
| - | Flowers 15-17 mm. Dorsal line sharp curved. | Orobanche pancicii f. curta |
| 5. | Corolla from base to edge mostly dark coloured. | 6. |
| - | Whole corolla dark coloured, often purple-black. | Orobanche pancicii f. melanochroa |
| 6. | Stem robust, up to 50 cm tall. Spike cylindrical, many-flowered and dense, later lax. Corolla 25-28 mm long, broadly campanulate. | Orobanche pancicii f. intumescens |
| - | Plant weakly. Spike few-flowered. Flowers very remote. Corolla tubular-campanulate. | Orobanche pancicii f. spaniantha |
Ic.: Beck, 1887 (tabl. VII fig. 8-12) Beck, 1890 (tabl. IV fig. 68) Beck, 1930 (p. 143 fig. 14 K) Pusch, 2000 (p. 41 f.) Király et al., 2005 (p. 20 fig. 1 and p. 23 fig. 3) Delipavlov, 1995 (p. 309 tabl. LXX fig. 3) Pusch, 2009 (p. 48 - male)
Maps: - Stoyanov, 2006 (p. 251 - Bulgaria) Pusch, 2009 (p. 49 - Europe) Dakskobler et al., 2010 (p. 172 - Slovenia)
Biology: - Fl. 5.6 (7). Meadows and pastures, in AUT edges of forests, meadows and pastures, road embankments, in BIH mesoxerophytic meadows and roadsides, in GRC also on alpine rocky slopes, cliffs, scree and meadows above the tree limit, on limestone and igneous rocks (granite), in SVN on supalpine (stony) grasslands and edges or clearings of pionerr coniferous forests (spruce or spruce-larch) on shallow soil (mainly rendzina) above limestone and dolomite; from foothills to the subalpine altitudinal range, (190) 240 - 1900 (2250) m, almost exclusively on Dipsacaceae.
Accompanying vegetation at locations in AUT: Achillea millefolium s.l., Aegopodium podagraria, Alopecurus pratensis, Arrhenatherum elatius, Campanula patula, Carex hirta, Cirsium oleraceum, Colchicum autumnale, Dactylis glomerata, Daucus carota, Equisetum arvense, Festuca pratensis, Fragraria viridis, Galium mollugo s.l., Heracleum sphondylium, Holcus lanatus, Knautia drymeia (host), Leucanthemum vulgare, Lilium martagon, Lotus corniculatus, Orobanche gracilis, Plantago lanceolata, Ranunculus acris, Rumex crispus, Trifolium pratense. Pusch, 2000 (p. 34 f.)
Accompanying vegetation at locations in BIH (Sarajevo): Briza media, Clinopodium vulgare, Colchicum autumnale, Dianthus deltoides, Euphorbia cyparissias, Filipendula vulgaris, Genista sagittalis, Gentiana cruciata, Geranium sanguineum, Hypericum hirsutum, Hypericum perfoliatum, Knautia arvensis, Linum catharticum, Lotus corniculatus, Malva moschata, Medicago falcata, Orobanche gracilis, Plantago media, Rhinanthus spec., Salvia verticillata, Scabiosa leucophylla (host), Stachys recta, Teucrium chamaedrys, Trifolium montanum. Pusch, 2000 (p. 35)
Accompanying vegetation at locations in BIH (Jahorina mts.): Achillea spec., Anemone narcissiflora, Campanula glomerata, Campanula patula, Epilobium angustifolium, Geranium sylvaticum, Geum rivale, Knautia dinarica (host), Linum capitatum, Lotus corniculatus, Myosotis spec. Orobanche gracilis, Phleum alpinum, Polygonum bistorta, Ranunculus croaticus, Salix silesiaca, Scabiosa ochroleuca, Solidago alpestris, Trifolium badium, Tussilago farfara, Valeriana officinalis. Pusch, 2000 (p. 35)
Host plants:
- Scabiosa leucophylla Borbas (Dipsacac.) sec. Beck, 1887 (p. 149) Beck, 1930 (p. 163) Hayek, 1928-31 (p. 225) Pusch, 2000 (p. 32) Pusch, 2009 (p. 49)
- Scabiosa lucida subsp. lucida Vill. (Dipsacac.) sec. Dakskobler et al., 2010 (p. 172 ff.)
- Knautia dinarica Maly (Dipsacac.) sec. Pusch, 2000 (p. 32)
- Knautia drymeia Heuff. (Dipsacac.) sec. Pusch, 2000 (p. 32) Király et al., 2005 (p. 22) Pusch, 2009 (p. 49) Dakskobler et al., 2010 (p. 172 ff.)
- Knautia dipsacifolia Kreutzer (Dipsacac.) sec. Beck, 1890 (p. 220 - O. reticulata f. viscosa) Pusch, 2009 (p. 49)
- Knautia fleischmannii (Hladnik ex Rchb.) Pacher (Dipsacac.) sec. Beck, 1930 (p. 163)
- Knautia longifolia (Waldstein & Kit.) Koch (Dipsacac.) sec. Dakskobler et al., 2010 (p. 172 ff.)
- Knautia magnifica Boiss. & Orph. ex Orph. (Dipsacac.) sec. E. Stamatiadou 1972 ATH 16035
- Knautia travnicensis G. Beck (Dipsacac.) sec. Benac, 1967 (p. 108 - only known from f. longebracteata)
- Cephalaria flava Szabó (Dipsacac.) sec. Stoyanov, 2006 (p. 251)
- Valeriana montana L. (Valerianac.) sec. Beck, 1930 (p. 163)
- Ligustrum vulgare L. (Oleac.) sec. Beck, 1887 (p. 149) Beck, 1930 (p. 163) Stojanov and Stefanov, 1948 (p. 1057) Stojanov et al., 1967 (p. 996) Pusch, 2009 (p. 49)
- Euonymus L. (Celastrac.) sec. Delipavlov, 1995 (p. 310) Kitanov and Penev, 1963 (p. 335) Stojanov and Stefanov, 1948 (p. 1057) Stojanov et al., 1967 (p. 996)
- Euonymus latifolia Mill. (Celastrac.) sec. Stoyanov, 2006 (p. 252)
The occurrence on Teucrium chamaedrys (Strid and Tan, 1991 p. 272) is erroneous. The parasitism on Pistacia terebinthus sec. Stoyanov, 2006 (p. 251) needs confirmation.
General distribution: - AUT, N ALB, N GRC, S BIH, SE and E SRB, BGR, E HUN, SVN and HRV. Actually also known from ITA. - Diagnoses: sm/mo.oz2 EUR E. Pusch, 2009 (p. 49) - Regional Arealdiagnoses: hell-balc-illyr-ostalpisch//mo-westpann. Pusch, 2009 (p. 49)
Examined specimen:
AUT: - Kärnten: Karawanken, N-Seite der Bärentaler Kotschna, vor der Plautzalm zum Gipfel, 1600-1914 m, subalp. staudenreiche Wiesen, MTB 9550/2, Schneeweiß 09.07.2003 WU 9712. - Kärnten: Karawanken, oberstes Bärental zwischen Narzissenwiese und Bärensattel, 1200-1698 m, mont. staudenreiche Wiesen, MTB 9550/2, Schneeweiß 11.07.2003 WU 9713. - Burgenland: 8964 Südliches Burgenland, SE Heiligenbrunn in einer kräuterreichen (bot. staudenreichen) Hangwiese zw. Weingärten reichlich auf Knautia drymeia, Helmut Melzer 23.05.2002 FR (H. Melzer: "Sehr starker Gewürznelkenduft! Merkwürdig, dass niemand gesagt hat, dass der Artname nicht korrekt ist, also nicht dem Code entspricht!") - Burgenland: 8964/4 ESE von Güssing bei Deutsch Bieling auf einer der letzten Bergwiesen (noch) zahlreich auf Knautia drymeia bei etwa 240 m Seehöhe, Helmut Melzer 08.06.2000 FR.
BIH: - "Ad radices Scabiosae leucophyllae, Bosn(ia): in montosis ca. Sarajevo", G. Beck (7) 06.1885 B, WU s.n. (Syntype) - Bosnia, ad radices Knautiarum in monte Romanja Pl(anina) ca. 1200 m, G. Beck 07.1888 JE, WU 214. - Abhänge des Trebevic bei Sarajevo, ca. 700 m, Franz Fiala 06.1894 WU s.n., det. Beck. - Hercegovina, bei Mostar, Padvelez, ca. 400 m, Egon Kindt 30.05.1905 WU s.n., det. Beck. - S-Hang des Vitorog Veliki, zwischen Büschen von Genista radiata, Kalk, 1700 m, J. Stadlmann, F. Faltis, E. Wibiral 10.07.1907 WU s.n., det. Beck
GRC: - North Central, Nomos Florinis: Ep. Florinis, Mt. Varnous (= Peristeri), 4 - 5 km N of Vigla, military post E of Bella vodha (= Kalo Nero; 40.48,5 N 21.13 E) summit, 1950 - 2100 m, alpine pasture and rocky slopes, igneous rocks (granite), mostly covered by Vaccinium myrtillus and Juniperus communis, parasitic on Knautia magnifica, E. Stamatiadou (25953) 25.06.1972 ATH 16035, det. Uhlich 14.03.1991; T. Raus & Kraft 21.08.1986 B 11213, 11214, det. Uhlich 02.09.1992. - North East, Nomos Serron/ Kavalas: Ep. Fillidos/ Pangeou, Mt. Pangeon, summit area, 1900 – 1950 m, limestone, A. Strid 14.07.1970 C 884, det. Hartvig 1989, confirm. Foley 03.1999; W. Greuter 15378, A. Strid 884 in Strid and Tan, 1991 p. 272 and in Karagiannakidou et al., 1995 p. 204. - North East, Nomos Dramas: Eparchia Dramas, Mt. Falakro, 5 - 8 km ESE of Volax, cliffs and scree in top area and meadows just above the tree-limit, 1550 – 2232 m, Gustavsson 1970 LD 67, 114.
Further cited locations:
HUN, Transdanubia: - Alpokalja, Köszegi-hegység: MTB 8665/1, Köszeg, Szabó-hegy, gesztenyésben (Visnaya 1933 BP s.n. as O. reticulata f. pallidiflora). Király et al., 2005 (p. 22) - Alpokalja, Köszegi-hegység: MTB 8665/1, Köszeg, a Szabó-hegy, É-i oldalan és a Kiraly-völggyel szomszédos gyertyános-tölgyesekben és gesztenyésekben sokfelé, összesen legalább, 390 m, on Knautia drymeia (Gergely Kiraly & A. Mesterházy 01.06.2005 BP s.n.; Gergely Kiraly 2005 in Herb. A. & G. Kiraly, Sopron; Z. Barina 2005 in BP). Király et al., 2005 (p. 22)
- Zalai-dombvidék, Kelet-dombság: MTB 9466/3, Bázakerettye (46.32 N 16.43 E), Kozari-erdö (Á. Károlyi 1945 BP; Á. Károlyi et al. 1971 as O. reticulata). Király et al., 2005 (p. 22) - Zalai-dombvidék, Kelet-dombság: MTB 9467/3, Obornak (46.28 N 16.52 E), in graminosis (Á. Károlyi 1948 BP; Á. Károlyi et al. 1971 as O. reticulata). Király et al., 2005 (p. 22) - Zalai-dombvidék, Kelet-dombság: MTB 9467/3, Homokkomárom (46.29 N 16.54 E), ad margines silvarum (Á. Károlyi 1949 BP; Á. Károlyi et al. 1971 as O. reticulata). Király et al., 2005 (p. 22) - Zalai-dombvidék, Kelet-dombság: MTB 9567/3, Szepetnek (46.26 N 16.55 E), a temetöben mintegy 25 tö (T. Horvath 2005 in Herb. A. & G. Kiraly, Sopron, det. Gergely Kiraly). Király et al., 2005 (p. 22)
AUT: - Niederösterreich: - on Knautia dipsacifolia, Saugraben of mt. Schneeberg, G. Beck 1882 or 1883 PRC s.n. Pusch, 2000 (p. 35 - loc. class. of O. reticulata f. viscosa)
- Steiermark: - E Steiermark, in vicinity of Klöch (N Bad Radkersburg): MTB 9261/2, Wald- Wiesen- und grabenränder am asphaltierten Fahrweg parallel zum Kutschenitza-Bach ca. 1 km N Pölten; 01.06.2000 (Hauptblüte ca. 14 Tage früher); ca. 19 Ex. auf Knautia drymeia, ca. 260 m s.m.! - Erstnachweis: Bregant & Maurer (1993) als "O. reticulata". Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - E Steiermark, in vicinity of Klöch (N Bad Radkersburg): MTB 9261/2, südexponierte Wegeböschungin Klöchberg (Hürtherberg) bei Klöch, ca. 60 m S Gasthof Palz; 01.06.2000 (Hauptblüte ca. 14 Tage früher); ca. 4 Ex. auf Knautia drymeia, ca. 280 m s.m.!- Erstnachweis: W. Maurer 1992 (Herbar W. Maurer), Bregant & Mauerer (1993) als "O. reticulata". Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - E Steiermark, in vicinity of Klöch (N Bad Radkersburg): MTB 9261/2, westexponierte Straßenböschung N Klöchan der Strasse Richtung Dt. Haseldorf (etwa am Nordende des Basaltsteinbruches); 01.06.2000 (Vollblüte); ca. 8 Ex. auf Knautia drymeia, ca. 360 m s.m.! - Erstnachweis. Pusch, 2000 (p. 38) - E Steiermark, in vicinity of Bierbaum (N Fürstenfeld): MTB 8962/1, Waldrand und Obstwiese ca. 600 m SW Bahnhof Bierbaum; 02.06.2000 (z.T. in Vollblüte); über 50 z.T. abgemähte Ex. auf Knautia drymeia, ca. 290 m s.m. - Erstnachweis: Heinrich 1923 in Fritsch 1926 als "O. reticulata var. viscosa", seither nicht wieder. Fritsch, 1926 (P. 223 as O. reticulata var. viscosa) Pusch, 2000 (p. 38) H. Uhlich obs. 29.07.2007 (few expl.) Pusch, 2009 (p. 49) - Eisenerzer Reichenstein, K. Tkalcsics in Pusch, 2009 (p. 49) - Schneealpe (at first L. Schratt-Ehrendorfer 2007). Pusch, 2009 (p. 49) - Koralpe. Pusch, 2009 (p. 49)
- Burgenland: - S Burgenland, in vicinity of Heiligenbrunn (E Güssing): MTB 8964/3 und 4, Wegränder und Obstwiesen im Kellerviertel (Kellergasse) in Heiligenbrunn, 01.06.2000 (Hauptblüte ca. 14 Tage früher); ca. 5 Ex. auf Knautia drymeia, ca. 240 m s.m.! - Erstnachweis: W. Maurer 1988 (Herbar W. Maurer). Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - Burgenland: S Burgenland, in vicinity of Heiligenbrunn (E Güssing): MTB 8964/4, Straßengraben an der Straße von Heiligenbrunn nach Strem etwa 700 m N Kirche Heiligenbrunn, 01.06.2000 (z.T. in Vollblüte), ca. 15 Ex. auf Knautia drymeia, ca. 240 m s.m.! - Erstnachweis! Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - Burgenland: S Burgenland, in vicinity of Heiligenbrunn (E Güssing): MTB 8964/3, Wald- und Bachrand ca. 600 m NNW Kirche Heiligenbrunn, 01.06.2000 (Vollblüte), 12 Ex. auf Knautia drymeia, hier gemeinsam mit Orobanche gracilis; ca. 270 m s.m.! - Erstnachweis! Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - Burgenland: S Burgenland, in vicinity of Heiligenbrunn (E Güssing): MTB 8964/4, Straßengräben und Wiesen ca. 1200 m und 1350 m SSE Kirche Heiligenbrunn (NW Deutsch Bieling), 05.06.2000, H. Melzer (briefl. Mitt.: insgesamt über 25 Ex. auf Knautia drymeia, bis auf 1 Ex. verblüht), ca. 240 m s.m. - Erstnachweis durch H. Melzer! Pusch, 2000 (p. 38) Pusch, 2009 (p. 49) - Burgenland: Gemeinde Allhau, Buchschachen, phot. J. Weinzettl 23.05.2004. Pusch, 2009 (p. 49) - Deutsch Bieling. Pusch, 2009 (p. 49)
- Carinthia: - Karawanken: Hochobir, ehemalige Bergwerkshalde unterhalb Eisenkappler Hütte in ca. 1500 m, auf Knautia drymeia, phot. K. Tkacsics 30.06.2002. Pusch, 2009 (p. 49) - Karawanken: Bärentaler Kotschna, einerseits nördlich unter dem Gipfel in 1750-1800 m, andererseits nordöstlich unter dem Bärensattel, ca. 1500 m, an beiden Stellen auf Knautia drymeia, G. M. Schneeweiß 09.07.2003 WU 9712, 9713. Pusch, 2009 (p. 49)
SVN: - 9547/4 (UTM 33TUM93) Julian Alps, the Loška stena ridge, under Mt. Morež, Zgornji Lepoč (Lašte) below the mountain pass Konjska škrbina, 1980 m, dolomite limestone, subalpine grassland, associated with Scabiosa lucida subsp. lucida and Knautia longifolia, I. Dakskobler 19.08.2008 ZRC SAZU (SRC SASA) 11595, conf. Pusch 27.02.2010. Dakskobler et al., 2010 (p. 172) - 9649/3 (UTM 33TVM13) Julian Alps, Bohinj, on the way from the alp Uskovnica towards Čiprje, near the hunting lodge below the peak Macesnova (Mesnova) glava, dolomite limestone, 1500 m, associated with Knautia drymeia, I. Dakskobler, B. Vreš, B. Anderle, I. Veber, B. Zupan & V. Leban 10.07.2008 ZRC SAZU (SRC SASA) 11594, conf. Pusch 27.02.2010. Dakskobler et al., 2010 (p. 172) - 9649/3 (UTM 33TVM13) Julian Alps, Bohinj, on the way from the alp Uskovnica towards Čiprje below the peak Močile, subalpine grassland north-east of the alp Trstje, dolomite limestone, 1500 m, associated with Scabiosa lucida subsp. lucida, I. Dakskobler, B. Vreš, B. Anderle, I. Veber, B. Zupan & V. Leban 10.07.2008, conf. Pusch 27.02.2010. Dakskobler et al., 2010 (p. 172 f.) - 9748/2 (UTM 33TVM02) Julian Alps, the lower Bohinj mts. (Spodnje Bohinjske gore), below the peak Konjski vrh (Konj) on the way to the Konjsko sedlo mtn. pass (under Mt. Bohinjski Migovec), 1530 m, dolomite limestone, subalpine stony grassland, associated with Scabiosa lucida subsp. stricta and Knautia longifolia, I. Dakskobler, I. Veber & B. Zupan 28.07.2008 ZRC SAZU (SRC SASA) 11593, conf. Pusch 27.02.2010. Dakskobler et al., 2010 (p. 173) - 9554/2 (UTM 33TVM84) Karavanke, the Peca ridge, below Mala Peca, 1550 m, limestone, forest edge, associated with Knautia drymeia, B. Vreš, I. Dakskobler, B. Anderle, V. Leban & V. Dakskobler 14.07.2009 ZRC SAZU (SRC SASA) 11590, conf. Pusch 27.02.2010. Dakskobler et al., 2010 (p. 173 f.) - S SVN: Mt. Travnik, phot. F. Müller s. a. (pers. comm.)
ITA: - Veneto: Prealpi Trevigiane, phot. Pino Perino (Picasa-user) 01.07.2009.
BIH: - Bosnia: - Ad rad. Scabiosae leucophyllae, Bosnia, in montosis circa Sarajevo, G. Beck (Plantae Bosniae et Hercegovinae, exsiccatae Nr. 7) 06.1885 WU s.n. Pusch, 2000 (p. 31) - Mt. Trebevic: "Magerrasen an einem asphaltiertem Fahrweg ca. 4,4 km SE Zentrum Sarajevo, auf Scabiosa leucophylla, am 24.06.2000, ca. 1130 m s.m. (auch G. Beck hat am 03.07.1888 'auf dem Trebevic' bei Sarajevo O. pancicii gesammelt. - Herbarbeleg in Prag PRC)". Pusch, 2000 (p. 32) - Jahorina mts.: "Fahrwegrand ca. 200 m NW Gipfel SE Zentrum Sarajevo, auf Knautia dinarica, am 24.06.2000, ca. 1840 m s.m.)." Pusch, 2000 (p. 32) - Veliki Vitorog, 1700 m, July (Stadlmann, Faltis, Wibiral). Beck, 1930 (p. 163) - Lisina mts. at Gornji Vakuf, 1300 m, July (K. Maly). Beck, 1930 (p. 163) - Mt. Vlasic at Travnik, 1400-1500 m, June (Beck). Beck, 1890 (p. 224) Vandas, 1891 (p. 253 as O. reticulata) Beck, 1930 (p. 163) - Vranica Planina on mt. Matorac (Beck). Beck, 1930 (p. 163) - At Zenica (Sendtner). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - On the Borova glava at Livno (Beck). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - Common in vicinity of Sarajevo (Beck, Fiala, K. Maly) an d on the nearest mts. Trebevic, Igman, Bjelasnica (-1760 m), Ozren, Romanija, Vratlo, Treskavica (-1700 m) (Beck, K. Maly). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - Lelija Planina at Jablan vrelo, 1550-1600 m, and on the Kablja Planina, ca. 1700 m, September (Vandas). Beck, 1930 (p. 163) - Mt. Maglic (Beck). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - Ljubnica (Curcic). Beck, 1930 (p. 163) - Mt. Stolac at Višegrad (Curcic). Beck, 1930 (p. 163)
- Herzegowina: - Valley of river Neretva between Raška gora and Vojno at 90 m (probably washed downwards), May (K.Maly). Beck, 1930 (p. 163) - Prenj mts., Prenj-Bjelašnica, Višocica (Beck). Beck, 1930 (p. 163) - Vran at Blidinje (Vandas). Beck, 1930 (p. 163) - Velez (Beck). Beck, 1930 (p. 163) - At Mostar (Fiala). Beck, 1930 (p. 163)
YUG: - Montenegro: at Livady, "In pascuis reg. subalp. sub monte Hum Orahovski distr. Kuci (Szyszylowicz)". Beck and Szyszylowicz, 1888 (p. 142) Beck, 1890 (p. 224) Beck, 1930 (p. 163) Rohlena, 1941-42 (p. 284) - SE Serbia: Stara Planina, Rakosch (Adamovic 1908). Josifovic, 1974 (p. 298) - E Serbia: Rtanj Planina prope Sako Banja (Pancic 1884 in Herb.Pancic). Beck, 1890 (p. 224) Formánek, 1895 (p. 62) Beck, 1930 (p. 163) Josifovic, 1974 (p. 298) - Montenegro: Mt. Volujak (Beck). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - SW Serbia: at Mrdzanov on mt. Zverda. Beck, 1890 (p. 224) Beck, 1930 (p. 163)
ALB: - N Albania: in vicinity of Pec at Bjelopolje, Bjelopolska Planina at 1350-1430 m, paras. on Valeriana montana. Beck, 1930 (p. 163) - At Vijenac. Beck, 1930 (p. 163) - Stavnjica dolnja ca. 1400 m and mt. Ples above Streoci at 1950 m, September, below Wulfenia (Vandas). Beck, 1930 (p. 163)
GRC: - Mts. of NE Greece. All known specimen examined by author (see above) - North East, Nomos Dramas: Eparchia Dramas, Mt. Falakro, 8 km ESE of Volax (= Volakas), cliffs and scree in top area and meadows just above the tree-limit, 41°18'23,45" N, 24°03'03,16" E, phot. A. Strid 28.07.1985 (Kit Tan pers. comm. 12.02.2012); phot. George Papagiannis 02.07.2011 on Hellenic Nature;
BGR: - Black Sea coast, Vitoschki, Sofijskij and Snepolski Rajon, from sea level up to 1800 m. Delipavlov, 1995 (p. 310)
- Vitosha region: - Vitoscha mts., W slope of mt. Kopitoto. Kitanov and Penev, 1963 (p. 335) - Vitoscha and Distr. Varna (Black Sea coast). Stojanov and Stefanov, 1948 (p. 1057) Stojanov et al., 1967 (p. 996) - Vitoša mts. (Skorpil, Velenovsky). Beck, 1890 (p. 224) Beck, 1930 (p. 163) - In declivibus graminosis m. Vitoša (Vel.). Velenovsky, 1891 (p. 440)
- West frontier Mts.: - Kyustendil province: Osogovska planina mts., meadows near Osogovo chalet. P. Hájková, 2003. Vegetation Science GroupMasaryk University Brno
- Western Rhodopes Mts.: - Peshtera, 750 m, K. Stoyanov 29.06.2005 SOA 57051. Stoyanov, 2006 (p. 251)
- Thracian plain: - On hills above Pazardzhik (Tatar Pazardzhik), 215 m, I. Mrkvichka 19.04.1914 SOA 48504. Stoyanov, 2006 (p. 251) - Asenovgrad, 300 m, on Pistacia terebinthus (?) and Cephalaria flava, K. Stoyanov 29.05.2003 SOA 57052 and 57053. Stoyanov, 2006 (p. 251) - Park Ayazmoto, 250 m, K. Stoyanov 01.06.2004 SOA 57054 and 57055. Stoyanov, 2006 (p. 251)
- Note: - According to Stoyanov, 2006 (p. 251) reported also for Nothern Black Sea Coast (Dimitrov, 2002; Stojanov and Stefanov, 1948; Stojanov et al., 1967; Delipavlov, 1995; Cheshmedzhiev, 2003; Kozhuharov, 1992 and Georgieff, 1937), Sofia Region (Dimitrov, 2002; Delipavlov, 1995; Cheshmedzhiev, 2003; Kozhuharov, 1992 and Urumov, 1912), Znepole Region (Dimitrov, 2002; Delipavlov, 1995; Cheshmedzhiev, 2003; Kozhuharov, 1992; Urumov, 1913a, Urumov, 1913b and Urumov, 1935b), Vitosha Region (Dimitrov, 2002; Stojanov et al., 1967; Delipavlov, 1995; Kitanov and Penev, 1963; Cheshmedzhiev, 2003; Kozhuharov, 1992; Georgieff, 1937; Urumov, 1930 and Velenovsky, 1891), Belasica Mtn. (Dimitrov, 2002; Cheshmedzhiev, 2003). It is indicated for: West Frontier Mts. (Urumov, 1913b, Urumov, 1935b), Struma Valley (Kozhuharov, 1992; Cheshmedzhiev, 2003; Dimitrov, 2002), Norther Struma Valley (Urumov, 1913a, Urumov, 1935b), Rila Mtn. (Urumov, 1935b), Strandsha Mtn.(Stojanov and Stefanov, 1948). The herbarium sheets and the collections of K. Stoyanov confirm the regions Northern Black Sea Coast, Sofia Region, Znepole Region, Vitosha Region, Belasica Mtn., Eastern Sredna Gora Mtns., Western Rhodopes Mtns. and Thracian Plain.
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, Kellergasse © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Heiligenbrunn, roadside between Heiligenbrunn and Deutsch Bieling © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Bierbaum a. d. Safen © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Bierbaum a. d. Safen © H. Uhlich 26.05.2011
Orobanche pancicii Burgenland: Bierbaum a. d. Safen © H. Uhlich 26.05.2011
